Decoyman
Bob-omb
I suspect the main thing they did was just touch up all assets while only redrawing very specific assets (+ in some cases altering a few animations overall to match some sprites with the modern games where they could, like Mario's shocked sprite).
For reference, most of Mario's sprites in TTYD were constructed using these assets. You can tell looking at the above image that Mario still uses the same torso from TTYD because unlike Origami King the buttons on his overalls aren't aligned perfectly horizontally (a specific quirk of TTYD's sprite interestingly), and they also use the exact same hand from TTYD. At best they've only been touched up a bit to suit the higher res, or they're being influenced by the new lighting engine. The Mario head is probably the only thing that was adjusted more specifically, more than likely to keep it consistent with the modern games because Paper Mario's actually gone through subtle expressive changes over the various entries (SPM has a much more tilted brim on his hat for example). The modern games have a lot of sprites where Mario faces the front, so I suspect they wanted this to be consistent with those games.
For reference, most of Mario's sprites in TTYD were constructed using these assets. You can tell looking at the above image that Mario still uses the same torso from TTYD because unlike Origami King the buttons on his overalls aren't aligned perfectly horizontally (a specific quirk of TTYD's sprite interestingly), and they also use the exact same hand from TTYD. At best they've only been touched up a bit to suit the higher res, or they're being influenced by the new lighting engine. The Mario head is probably the only thing that was adjusted more specifically, more than likely to keep it consistent with the modern games because Paper Mario's actually gone through subtle expressive changes over the various entries (SPM has a much more tilted brim on his hat for example). The modern games have a lot of sprites where Mario faces the front, so I suspect they wanted this to be consistent with those games.
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